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"I like them!" ~Willie's only line in Madagascar
"WE'RE ALL GONNA DIEEEE!" ~Willie's signature phrase in All Hail King Julien
Willie's first appearance was in the episode, "Poll Position," in which he is told that he is to become the captain of King Julien's first navy ship named Old Porceleinesides. He appears to be slightly smarter then the other lemurs, as he was imminently distraught by the fact that the ship he was to captain didn't look very seaworthy. Still with some motivation from Maurice (who poked him with a stick to get him to move towards the boat) he gave in and accepted the honor. His worries about the boat being unseaworthy are confirmed later in the episode when the boat starts sinking, a few minutes later after King Julien flipped the switch to release the boat into the water. He is later seen in the episode being attacked by very aggressive dolphins while the boat is sinking. When he is angry with King Julien for putting him in this predicament (the sinking ship), he lets himself be bribed with the mango's king Julien was sharing to increase his popularity. After catching the mango he screams your alright King J. He is either very lucky, able to swim or rescued from the water by Clover, as he appears in the next episode in which he celebrates Franksgiving in the honor of the sky god named Frank. He once tried to smuggle in more then one wishing stone in the wishing balloon something which he is called out for by Maurice.
Willie is a very hyper and fearful lemur. He can however be very tough and aggressive if he's armed with dangerous weapons. Despite being slightly smarter than many of the other lemurs, he has a lot of immature moments, such as when he mischievously called out Dorothy for not wearing a diaper, and such as when he keeps screaming, "WE'RE ALL GONNA DIE!" When he screams his signature phrase, he often causes a huge panic among the citizens of the kingdom; he will however sometimes scream this when panic is already happening. Like a lot of the others, however, Willie is very fun-loving.