Penguins of Madagascar is a computer-animated adventure comedy film directed and produced by DreamWorks Animation, distributed by 20th Century Fox. Based on related TV series The Penguins of Madagascar, the film was released on November 26th, 2014.
Skipper, Kowalski, Rico, and Private are brought into the services of the North Wind, an Arctic elite undercover inter-species task force that is dedicated to helping animals who can't help themselves. The Penguins must work with the North Wind led by Classified in order to prevent Dr. Octavius Brine from taking over the world.
The move begins in Antarctica. Multiple penguins are walking somewhere, while baby Skipper, Kowalski, and Rico ask where they are even going - but the other penguins. don't seem to care. And when an egg rolls by, they comment that they lose a few eggs, and it's nature. Skipper decides to go against nature, and goes after the egg which lands safely on a boat... full of Leopard Seals.
Rico eats the egg and they fool the seals into releasing the harpoon and sending them onto a drifting iceberg. The egg hatches, revealing a baby Private. Private ask if the trio are his family. Kowalski tells him that he doesn't have a family they will die, Skipper slaps Kowalski and tells Private that they are his family now. The iceberg drifts away to where adventure will lead them.
The scene moves on to where Madagascar 3: Europe's Most Wanted left off. The penguins (except for Private) have grown tired of Marty's Afro Circus song along with King Julien's song I Like to Move It, Move It and plan to take a break from the circus by breaking into the United States gold depository to celebrate a 10 year old private's birthday. After they infiltrate the building, Skipper asks Private what he would in the world. Private responds by saying he would be a meaningful and valued member of the team but Skipper says they got him something else - Cheezy Dibbles. When Private tries to get a bag the vending machine pulls him in. The penguins try to get him out but it grabs them all, bursts out the building and is carried from a helicopter to a submarine.
The penguins escape the vending machine later, having eaten all of the Cheezy Dibbles, and escape a cage using Rico's paperclip and Skipper calls him a meaningful and valued member of the team. On the control console, Private starts pressing buttons, which causes them to move down levels. They encounter a human who strangely knows their names. Skipper ask who he is, and he responds by saying he is Doctor Octavirus Brine but he is really an octopus named Dave - but none of the penguins remember. Dave explains that he used to be a star attraction at Central Park Zoo, until the penguins arrived. Dave was moved from zoo to zoo, always being upstaged by penguins. He shows a substance he created and orders his octapi to attack the penguins. The penguins manage to steal the substance and escape into the canals of Venice, going on a canoe chase to escape. However, they are cornered.
Luckily they are rescued - a jet appears and a wolf tells them they are now under the protection of the North Wind. Inside the jet Skipper demands answers. The wolf says that the North Wind is an elite undercover force devoted to helping animals who can't help themselves. They introduce themselves as Eva, Short Fuse, and Corporal, and Skipper refers to the wolf as Classified. Skipper proudly introduces his own team, but the North Wind is less than impressed, and returns to their headquarters.
Skipper is impressed with their technology. Classified wants the penguins to tell him everything they know about Dave. Skipper tells him that Dave is an octopus, and that they managed to steal the Medusa Serum, but then Dave calls the North Wind. Classified mocks Dave for the loss of his serum, but Dave reveals that he has a huge tank of it and reveals his plan is to use it on all penguins everywhere. The Penguins get ready to go on the mission with Dave, but he is still convinced that they are useless, and has them tranquilised and sent to a safe house.
The penguins wake up in a cargo plane to Madagascar. Skipper and Kowalski realize what's happening and escape the plane. After crashing to the ground they begin to wonder where Dave will strike next. Meanwhile in Rio Dave kidnaps more penguins and leaves just as the North Wind arrive. Corporal begins to have his stress eating again, while Eva picks up the the penguins never arrive to Madagascar.
The penguins arrive at Shanghai (which they mistake for Dublin) and thanks to Rico hacking up Dave's snowglobe collection, realise that Dave is going to strike Shanghai. In the zoo, Skipper has Private create the diversion, dressed as a mermaid. Dave arrives, but is captured by the Penguins when the North Wind arrive and want to take him in, but Dave escapes into the pipes, and captures the mermaid penguins, and Private with them. He escapes to his submarine, and the Penguins hijack the North Wind's jet, while the North Wind tries to catch them. Ultimately the Penguins can't fly the jet properly, and the jet explodes, with Dave getting away. The Penguins are tied adrift from the North Wind's life-boat. It turns out the North Wind has been tracking Private from his tranquilizer dart and finds the land Dave resided his sub in.
On the island hudreds of penguins are being held captive, and see Dave's demonstration of the Medusa serum to turn them into hideous monsters. Private accidentally blows his cover, prompting Dave to use him as his next victim.
Outside, the penguins and North Wind debate whose plan to use - and after Classified reminds Skipper that it's his fault Private was captured, he agrees to Dave's plan, while the Penguins cause a diversion - however, Dave was prepared for their arrival and captures first the North Wind, then the Penguins.
Dave reveals that he will use Private as his first Test subject. The penguins try to get him out, but can't find Rico's paperclip, and Private is disintegrated. Though the penguins are dismayed, it turns out Private escaped the ray with Rico's paperclip. First he rescues the North Wind from Dave's complicated death device. Private wants to go save the other penguins, but Classified tells him that they have to go back to base and get new gear first. Private disagrees, and heads back to rescue the others.
But he is too late - Dave mutates the Penguins, then unleashes them on Manhattan. The exterminators arrive en masse, while Private manages to find Skipper, and talks sense into him. They find Kowalski, and then Rico, who also remember him, although Skipper admits Rico is much the same either way. The brothers set out to save the peinguins, and Private believes that they can reverse the ray.
But Kowalski says that they need to replace the ray with something extremely cute. They see Private hooking himself up the ray and protest, but Private is adamant. Meanwhile Dave is realising that his revenge hasn't made him feel any better - so he decides to start mutating everything that's cute. After seeing the penguins with his ray he has his octopi capture them. The Penguins use Dave's snow globe collection to amplify and mirror the blast, but the batteries are dead and the battle resumes.
Rico is sent to go find batteries while they take on the Octopi. Rico finds batteries but is distracted by Cheezy Dibbles. The penguins realize they cannot hold them off much longer, luckily the North Wind show up in an Ice Cream van which explodes upon impact on Dave's ship. The North Wind walk away from the explosions before being sucked up by an exterminators vehicle. Dave fires himself at the cannon and throws Kowalski at the exterminator's vehicle. Dave tells Skipper he's lost but Skipper points out that Rico is tossing him double AAs. Skipper puts the Batteries inside but Dave tosses him and Rico at the exterminators vehicle before going after Private. Private uses his butt hand to punch Dave away. Skipper uses a dibble to aim for the button he succeeds before being sucked in, activating the ray and curing the penguins.
However, Private is overwhelmed by a blast mutates rapidly before a chrysalis forms around him. Corporal manages to get him out and suddenly it haches turning Private magenta purple with spots and antlers. The group shares a hug. Skipper tells Private that he is a meaningful and valued member of the team, and everybody cheers him - then it turns out Dave who had taken most of the effect on the ray has shrunk and somehow became trapped in a snow globe. The penguins give him to to a little girl who shakes him up and down.
Classified apologizes for his speciesist attitude towards the penguins and wants to make it up. After a romantic moment between Kowalski and Eva, Private suggest jetpacks which everyone agrees on. The penguins are seen flying back to the circus, and it is implied they are now members of the North Wind.
In the epilogue Private asks Skipper if he will turn him normal and Skipper agrees, this time using Mort as the source of cuteness. They turn Private normal again. Mort doesn't seem to suffer any changes - until he eats King Julien in one gulp.
- Tom McGrath as Skipper, the leader of the penguins
- Chris Miller as Kowalski, the brains of the penguins
- Conrad Vernon as Rico, the loose cannon of the penguins*
- Christopher Knights as Private, the rookie of the penguins
- Benedict Cumberbatch as Classified, a wolf and team leader of the North Wind.
- John Malkovich as Dave, a disgruntled octopus who poses as the human Dr. Octavius Brine.
- Ken Jeong as Short Fuse, an explosive and demolitions expert seal and a member of the North Wind.
- Annet Mahendru as Eva, a snowy owl and the North Wind's intelligence analyst.
- Peter Stormare as Corporal, a Norwegian polar bear and a member of the North Wind.
- Werner Herzog as Documentary Filmmaker
- Danny Jacobs as King Julien
- Andy Richter as Mort